/* Edit the palette and sizing here. No build tools or external fonts required. */
:root{--paper:#f7f5ee;--ink:#203e35;--body:#45534b;--muted:#606a60;--line:#d4d9ce;--green:#e8eddf;--white:#fffef9;--accent:#acbb83;--serif:Georgia,"Times New Roman",serif;--sans:-apple-system,BlinkMacSystemFont,"Segoe UI",sans-serif}

*{box-sizing:border-box}
html{scroll-behavior:smooth;scroll-padding-top:28px}
body{margin:0;background:var(--paper);color:var(--body);font:16px/1.7 var(--sans);-webkit-font-smoothing:antialiased}
a{color:inherit;text-underline-offset:5px;text-decoration-thickness:1px}
a:hover{color:#122e22}
a:focus-visible,summary:focus-visible{outline:3px solid #536d2b;outline-offset:6px;border-radius:2px}
img{max-width:100%;display:block}
p,h1,h2,h3,figure{margin:0}
p+p{margin-top:16px}
h1,h2,h3{color:var(--ink);font-weight:400}
h1,h2{font-family:var(--serif);line-height:1.08;letter-spacing:-.045em}
h1{font-size:clamp(52px,5.7vw,82px)}
h1 em{font-weight:400;color:#607349}
h2{font-size:clamp(34px,3.5vw,49px)}
h3{line-height:1.3;letter-spacing:-.025em}
.wrap{width:min(1184px,calc(100% - 96px));margin-inline:auto}
.skip-link{position:absolute;top:-100px;left:20px;background:var(--ink);color:white;padding:12px 20px;z-index:10}
.skip-link:focus{top:12px;color:white}
.site-header{min-height:106px;display:flex;align-items:center;justify-content:space-between;border-bottom:1px solid var(--line);gap:24px}
.wordmark{text-decoration:none;font-size:22px;font-weight:650;letter-spacing:-.9px;color:var(--ink);white-space:nowrap}
.wordmark-dot{color:#7c914a;margin-left:2px}
nav{display:flex;gap:29px;align-items:center;font-size:14px}
nav a{text-decoration:none;padding-block:10px}
nav a:hover,nav a[aria-current]{text-decoration:underline}
nav .nav-contact{border:1px solid #bac3b3;border-radius:30px;padding:9px 17px}
.nav-contact span{margin-left:12px}
.hero{display:grid;grid-template-columns:1.2fr 1fr;align-items:center;gap:26px;padding-block:85px 68px}
.eyebrow{text-transform:uppercase;letter-spacing:.12em;font-size:11px;font-weight:650;line-height:1.6;color:var(--muted)}
.status-dot{display:inline-block;width:7px;height:7px;border-radius:100%;background:#728548;margin:0 8px 1px 0}
.hero h1{margin:25px 0 26px}
.hero-intro{font-size:19px;line-height:1.65;max-width:480px}
.hero-affiliation{font-size:13px;color:var(--muted);margin-top:25px;line-height:1.7}
.actions{display:flex;align-items:center;gap:26px;margin-top:31px}
.button{display:inline-flex;align-items:center;justify-content:center;gap:25px;background:var(--ink);color:var(--white);text-decoration:none;font-size:13px;border-radius:5px;padding:13px 20px;min-height:48px}
.button:hover{background:#315b49;color:white}
.text-link{display:inline-flex;gap:11px;align-items:center;min-height:44px;font-size:14px;font-weight:550}
.file-type{font:10px var(--sans);border:1px solid #b4beae;padding:2px 4px;text-decoration:none}
.hero-art{align-self:center;text-align:center}
.hero-art img{width:100%;height:auto}
.hero-art figcaption{font-size:11px;color:var(--muted);line-height:1.8;margin-top:-15px}
.figure-number{font-size:9px;margin-right:10px;letter-spacing:.1em}
.caption-note{color:var(--muted);font-size:10px}
.research-strip{display:flex;align-items:center;gap:18px;padding-block:22px;border-block:1px solid var(--line);font-size:11px;text-transform:uppercase;letter-spacing:.1em;color:var(--muted)}
.strip-line{height:1px;background:#a0ac98;width:60px}
.strip-index{margin-left:auto}
.section{padding-block:85px}
.section-heading{display:flex;justify-content:space-between;align-items:flex-end;gap:36px;margin-bottom:38px}
.section-heading h2{margin-top:16px}
.section-heading>p{font-size:13px;line-height:1.8;color:var(--muted);padding-bottom:3px}
.project{display:grid;grid-template-columns:220px 1fr;gap:40px;padding:40px;border:1px solid #ccd5c3;border-radius:8px}
.project-featured{background:var(--green)}
.project+.project{margin-top:22px}
.project-secondary{background:transparent}
.project-label{display:flex;align-items:flex-start;flex-direction:column;gap:20px}
.project-number{font:italic 44px var(--serif);color:#7d8c70}
.pill{font-size:10px;text-transform:uppercase;letter-spacing:.08em;border:1px solid #a7b498;padding:5px 9px;border-radius:24px;white-space:nowrap;color:#4c6042}
.project h3{font:36px/1.14 var(--serif);letter-spacing:-.03em;margin:14px 0 22px}
.project-content>p:not(.eyebrow){max-width:695px}
.workflow{display:grid;grid-template-columns:repeat(3,1fr);border-block:1px solid #c3ceba;margin:26px 0 12px;padding-block:17px;gap:16px;font-size:12px}
.workflow span{display:block;font:11px var(--sans);color:#657658;margin-bottom:7px}
.tags{display:flex;gap:8px;flex-wrap:wrap;margin-top:20px}
.tags span{font-size:11px;padding:4px 9px;border-radius:4px;background:#f7f8f166;border:1px solid #c9d2c1;color:#52624a}
details{margin-top:20px}
summary{display:flex;align-items:center;justify-content:space-between;cursor:pointer;list-style:none;padding:12px 0;border-bottom:1px solid #c5cdbf;font-size:13px;font-weight:600;color:var(--ink);gap:20px}
summary::-webkit-details-marker{display:none}
summary span{font-size:20px;font-weight:400;transition:transform .2s}
details[open] summary span{transform:rotate(45deg)}
.detail-content{font-size:14px}
.detail-content ul{padding-left:20px;margin:20px 0}
.detail-content li+li{margin-top:10px}
.cell-art{position:relative;width:137px;height:137px;background:#e5e8da;border:1px solid #aab89b;border-radius:47% 53% 58% 42% / 47% 46% 54% 53%;margin:22px 0 0 10px;box-shadow:inset 0 0 0 10px var(--paper),inset 0 0 0 11px #b9c4ab;transform:rotate(-12deg)}
.cell-nucleus{position:absolute;width:56px;height:64px;border:1px solid #91a480;background:#bac7a8;border-radius:55% 45% 39% 61%;left:34px;top:36px}
.cell-art i{position:absolute;width:9px;height:6px;background:#9caf83;border-radius:50%;top:29px;left:70px}
.cell-art i:nth-of-type(2){top:87px;left:104px}
.cell-art i:nth-of-type(3){top:103px;left:45px}
.publications-section{background:#eeeee4;border-block:1px solid var(--line)}
.publication-list{list-style:none;margin:0;padding:0}
.publication-list li{display:grid;grid-template-columns:110px 1fr;gap:35px;padding:29px 0;border-top:1px solid #cbd0c2}
.pub-year{font:italic 25px var(--serif);color:#69775e}
.publication-list h3{font:25px/1.35 var(--serif);max-width:900px;margin:10px 0 11px}
.publication-list h3 a{text-decoration:none}
.publication-list h3 a:hover{text-decoration:underline}
.publication-list h3 span{font:20px var(--sans);white-space:nowrap}
.publication-list p:not(.eyebrow){font-size:12px;color:var(--muted)}
.publication-list strong{font-weight:600;color:var(--body)}
.manuscript{display:flex;align-items:flex-start;gap:26px;border-top:1px solid #cbd0c2;padding-top:24px;margin-top:5px}
.manuscript p{font-size:13px}
.manuscript p span{font-size:11px;color:var(--muted)}
.about-section{display:grid;grid-template-columns:1fr 2fr;gap:80px}
.about-section h2{margin:17px 0 24px}
.large-copy{font:27px/1.45 var(--serif);color:var(--ink);letter-spacing:-.025em}
.about-copy>p+p{margin-top:22px}
.about-grid{display:grid;grid-template-columns:1fr 1fr;gap:32px;margin:32px 0;padding-top:27px;border-top:1px solid var(--line)}
.about-grid h3,.expertise h3{font-size:13px;font-weight:650;margin-bottom:12px}
.about-grid p{font-size:13px;line-height:1.8}
.about-grid strong{font-weight:600}
.expertise{border-top:1px solid var(--line);padding-top:25px}
.expertise p{font-size:13px;line-height:1.9}
.thoughts-teaser{display:flex;align-items:center;justify-content:space-between;gap:25px;padding:35px 0 50px;border-top:1px solid var(--line);margin-bottom:40px}
.thoughts-teaser h2{font-size:36px;margin:14px 0}
.thoughts-teaser p:not(.eyebrow){font-size:14px}
.thoughts-teaser .text-link{white-space:nowrap}
.contact-section{background:var(--ink);color:#e2e8d9}
.contact-inner{padding-block:68px;display:grid;grid-template-columns:1.4fr 1fr;align-items:center;gap:40px}
.contact-section .eyebrow{color:#c1cdaa}
.contact-section h2{color:var(--paper);font-size:45px;margin:18px 0 24px}
.contact-section p:not(.eyebrow){font-size:13px;color:#c5d0c3}
.contact-links{display:flex;align-items:flex-start;flex-direction:column;gap:16px;font-size:14px}
.contact-links a{min-height:28px}
.contact-links a:hover{color:white}
.contact-links a:focus-visible{outline-color:#d4e5ad}
.contact-links span{margin-left:12px}
.contact-links .email-link{font-size:18px;padding-bottom:15px;border-bottom:1px solid #5f7468;overflow-wrap:anywhere}
.site-footer{display:flex;justify-content:space-between;align-items:center;gap:20px;padding-block:25px;font-size:10px;color:var(--muted)}
.site-footer p+p{margin:0}
.site-footer a{padding-block:10px}
.thoughts-main{min-height:calc(100vh - 190px);padding-block:80px}
.thoughts-header{max-width:740px}
.thoughts-header h1{margin:20px 0 28px}
.thoughts-header .large-copy{max-width:600px}
.empty-notebook{margin-top:60px;padding:38px;border:1px solid var(--line);border-radius:8px;display:flex;gap:40px;background:var(--green);max-width:850px}
.notebook-mark{font:64px var(--serif);color:#7b905c}
.empty-notebook h2{font-size:32px;margin:12px 0}
.empty-notebook p:not(.eyebrow){max-width:500px}
.empty-notebook .text-link{margin-top:20px}
.thought-entry{max-width:760px;border-top:1px solid var(--line);margin-top:50px;padding-top:30px}
.thought-entry h2{font-size:34px;margin:15px 0 22px}
.thought-entry time{font-size:13px;color:var(--muted)}

@media(min-width:1450px){.hero{padding-block:100px 85px}
}

@media(max-width:1000px){.wrap{width:calc(100% - 56px)}
nav{gap:18px}
.project{grid-template-columns:155px 1fr;gap:25px;padding:30px}
.project h3{font-size:32px}
.hero{gap:0}
.hero-intro{font-size:17px}
.actions{gap:16px}
.about-section{gap:40px}
.contact-links .email-link{font-size:16px}
.desktop-break{display:none}
.section-heading>p{max-width:240px}
.strip-index{display:none}
}

@media(max-width:760px){.site-header{flex-direction:column;align-items:flex-start;gap:10px;padding-block:20px;min-height:0}
.wordmark{font-size:23px}
nav{width:100%;justify-content:space-between;gap:10px;font-size:12px}
nav .nav-contact{border:0;padding:10px 0}
.nav-contact span{display:none}
.hero{grid-template-columns:1fr;padding:48px 0 37px}
.hero-copy{max-width:560px}
.hero h1{font-size:clamp(49px,9vw,70px);margin:20px 0}
.hero-intro{font-size:18px}
.hero-art{max-width:390px;justify-self:center;margin-top:25px}
.hero-art figcaption{margin-top:-20px}
.hero-art img{max-height:335px}
.section{padding-block:52px}
.section-heading{display:block;margin-bottom:26px}
.section-heading>p{margin-top:20px;max-width:none}
.section-heading h2{font-size:37px}
.project{grid-template-columns:1fr;gap:21px;padding:25px}
.project-label{flex-direction:row;align-items:center;gap:20px}
.project-number{font-size:32px}
.cell-art{display:none}
.project h3{font-size:32px}
.project-content>p:not(.eyebrow){font-size:15px}
.project-content .eyebrow{font-size:10px}
.workflow{font-size:11px;gap:9px}
.research-strip{font-size:9px;gap:12px}
.strip-line{width:30px}
.publication-list li{grid-template-columns:48px 1fr;gap:15px;padding-block:24px}
.publication-list h3{font-size:23px}
.pub-year{font-size:20px}
.manuscript{display:block}
.manuscript p{margin-top:13px}
.about-section{grid-template-columns:1fr;gap:25px}
.about-section h2 br{display:none}
.about-section h2{margin-bottom:15px}
.large-copy{font-size:25px}
.about-grid{gap:24px}
.thoughts-teaser{display:block;padding-top:30px;margin-bottom:20px}
.thoughts-teaser .text-link{margin-top:15px}
.contact-inner{grid-template-columns:1fr;gap:32px;padding-block:45px}
.contact-section h2{font-size:39px}
.contact-links .email-link{font-size:17px}
.site-footer{flex-wrap:wrap;gap:10px;font-size:10px}
.site-footer p:nth-child(2){display:none}
.thoughts-main{padding-block:45px}
.empty-notebook{margin-top:40px;padding:26px;gap:20px}
.notebook-mark{font-size:38px}
.empty-notebook h2{font-size:28px}
}

@media(max-width:390px){.wrap{width:calc(100% - 36px)}
nav{font-size:11px;gap:8px}
.actions{gap:16px;flex-wrap:wrap}
.button{padding-inline:16px}
.hero h1{font-size:47px}
.eyebrow{font-size:10px}
.project{padding:20px}
.about-grid{grid-template-columns:1fr}
.empty-notebook{display:block}
.notebook-mark{display:block;margin-bottom:16px}
}

@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}
*,*::before,*::after{transition:none!important}
}

@media print{body{background:white;color:black;font-size:11pt}
.wrap{width:100%}
.site-header nav,.hero-art,.thoughts-teaser,.site-footer,.actions,.research-strip,.cell-art{display:none}
.site-header{min-height:50px}
.hero,.section{padding-block:25px}
.hero,.about-section,.contact-inner{display:block}
h1{font-size:36pt}
h2{font-size:25pt}
.project{display:block;padding:20px;margin-top:20px;break-inside:avoid}
.project-label{flex-direction:row;margin-bottom:15px}
.contact-section{background:white;color:black}
.contact-section h2,.contact-section .eyebrow,.contact-section p:not(.eyebrow){color:black}
.contact-inner{padding:25px 0}
.publications-section{background:white}
.publication-list li{break-inside:avoid}
.contact-links{margin-top:15px}
.hero-affiliation{margin-top:10px}
}

/* Keep navigation and contact targets comfortable on touchscreens. */
nav a{min-height:44px;display:inline-flex;align-items:center}

.contact-links a,.site-footer a{min-height:44px;display:inline-flex;align-items:center}

.hero-copy,.project-content,.about-copy{min-width:0}